Howe's record in the war was marked by the costly assault on Breed's Hill known as the Battle of Bunker Hill and the successful capture of New York City and Philadelphia - the latter of which would have significant strategic implications.< William was born in England, the third son of Emanuel Howe, 2nd Viscount Howe and Charlotte, the daughter of Sophia von Kielmansegg, Countess of Leinster and Darlington - a half-sister of King George I. This connection with the crown may have improved the careers of all three sons, but all were also very capable officers.

William's eldest brother was General George Howe, who was killed at Ticonderoga in 1758. His next brother was Admiral Richard Howe, who joined him in America during the revolution. He entered the army when he was seventeen by buying a Cornet's commission in the Duke of Cumberland's Dragoons in 1746. By the next year, he was fighting as a Lieutenant in Flanders as a part of the War of the Austrian Succession. After this war, he joined the 20th Regiment of Foot where he became a friend of James Wolfe.
During the Seven Years' War, Howe's service first brought him to America. His service in this conflict did much to raise his reputation. William commanded a regiment at the siege of Louisbourg and led a successful amphibious landing. This action, carried out under fire, won the attackers a flanking position and earned Howe his commander's praise.Howe commanded the light infantry under Major General James Wolfe at the Battle of Quebec, Canada on September 13, 1759. He led a fighting ascent to gain position on the Plains of Abraham, clearing the way for Wolfe's army to assemble before that battle.< His actions here earned him the rank of Brigadier General. He earned further fame in the capture of Montreal under Jeffrey Amherst before returning to England.
Howe also served in the capture of Belle Isle, off the French coast, in 1761. He was adjutant-general of the force that captured Havana in 1762. In 1772, Howe was elected a Member of Parliament for Nottingham. This was not unusual, as the election of 1761 sent more than 60 army officers to the British House of Commons. He was generally sympathetic to the American colonies. He opposed the Coercive Acts, and, in 1774, assured his constituents that he would resist active duty against the Americans.
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